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Divergent stereoview of the structure of metal-free (apo) EhDbr1. The σA-weighted 2Fo-Fc (light blue mesh) is contoured at 1.2 sigma. In the absence of metals, a water binds in the β-pocket, and a sulfate from the crystallization solution hydrogen bonds to the catalytic acid H91. His16 adopts an “out” position, rotating away from the position where it could stack with a branchpoint adenosine and ligate the α-pocket metal. Hydrogen bonds are dashed lines.
