A hypothetic mechanistic model for Pms1–regulated PSMS in rice. The PMS1T transcript is targeted by miR2118, and more 21-nt phasiRNAs are produced in 58S than MH63 under long-day conditions because of the SNP S2 (red dot as base T, black dot as base G). The accumulation of phasiRNAs eventually causes male sterility in rice. The SNP S2 may influence the cleavage efficiency mediated by miR2118, resulting in more PMS1T-phasiRNAs in 58S under long days. On the other hand, PMS1T-phasiRNAs may function as secondary small RNA to target some unknown transcripts, which should play important roles in male sterility in rice.