Table 1. Generalized Linear Mixed Model of overall performance in the honey tool-task.

	B	SE	Z	p value
(Intercept)	−2.588	1.33	−1.943	0.052
Human Orientation Index	0.227	0.08	2.699	0.007**
Novel food: time until touch	−0.034	0.01	−2.263	0.024*
Novel object: time until touch	−0.025	0.01	−2.123	0.034*
Novel object: exploration duration	0.003	0.01	0.433	0.665
Age	−0.006	0.03	−0.232	0.817
SEX (male)	0.124	0.51	0.243	0.808
Species (Sumatra)	−0.094	0.59	−0.16	0.873
Background
Wild vs. Rest	−0.173	0.27	−0.639	0.523
Rehab vs. Zoo	−0.402	0.29	−1.395	0.163
Unknown vs. Rehab.Rest	0.073	0.20	0.367	0.714
Human vs. Station (within rehabilitation station)	−0.479	0.48	−1.008	0.314
Mother vs. Hand (within zoo)	0.365	0.51	0.722	0.470
Accessibility (trend analysis)
Linear	0.786	0.76	1.04	0.298
Quadratic	0.228	0.89	0.257	0.797
Cubic	−0.189	0.89	−0.213	0.831
Sub-task (trend analysis)
Linear	−4.969	0.74	−6.737	<0.001***
Quadratic	1.554	0.44	3.559	<0.001***
Cubic	−0.205	0.33	−0.626	0.531

Note: The model is controlling for repeated observations on each facility and individual. The performance in the honey tool-task was binary measured. The Analysis included 88 individuals in 9 different zoos/rehab stations, totalling 352 observations, χ² = 226.27, P < 0.001, P-values below 0.05 appear in bold. Parameter estimates from a binomial GLMM, predicting the probability of an animal solving the task.