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. 2004 Oct;136(2):3350–3363. doi: 10.1104/pp.104.040758

Table I.

Symbols used in text

A Net photosynthesis rate
ā Weighted mean discrimination against C18OO for diffusion from chloroplast to atmosphere
a Discrimination against C18OO during diffusion through stomata
ab Discrimination against C18OO during diffusion through leaf boundary layer
aw Summed discriminations against C18OO during liquid phase diffusion and dissolution
a13 Discrimination against 13CO2 during diffusion through stomata
ab13 Discrimination against 13CO2 during diffusion through leaf boundary layer
aw13 Summed discrimination against 13CO2 during dissolution and liquid phase diffusion
αw Equilibrium oxygen isotope effect between CO2 and water
b Discrimination against 13CO2 by carboxylating enzymes
b18 Discrimination against C18OO by Rubisco
C Molar concentration of water
Ca Partial pressure of CO2 in atmosphere
Cc Partial pressure of CO2 in chloroplast
Ccs Partial pressure of CO2 at the chloroplast surface
Ci Partial pressure of CO2 in leaf intercellular air spaces
Cin Partial pressure of CO2 in dry air entering leaf chamber
Cs Partial pressure of CO2 at the leaf surface
ci Mole fraction of CO2 in intercellular air spaces
ca Mole fraction of CO2 in atmosphere
D Diffusivity of H218O in water
ΔA Discrimination against 13C or 18O during net CO2 uptake by photosynthesis
Δca 18O enrichment of CO2 in chloroplast compared to atmosphere
Δe 18O enrichment at evaporative sites in leaves compared to source water
Δea 18O enrichment of CO2 in chloroplast compared to atmosphere when chloroplast CO2 is in full equilibrium with chloroplast water
Δi Discrimination against 13CO2 that would occur if gi were infinite and photorespiration and day respiration did not discriminate
ΔL 18O enrichment of average lamina leaf water compared to source water
Δobs Observed discrimination against 13CO2 during photosynthesis
Δv 18O enrichment of vapor in atmosphere compared to source water
δA δ18O of CO2 taken up by net photosynthesis (VSMOW scale)
δa δ18O of CO2 in atmosphere (VSMOW scale)
δc δ18O of CO2 in chloroplast (VSMOW scale)
δc0 δ18O of CO2 in chloroplast that has not equilibrated with chloroplast water (VSMOW scale)
δe δ18O of water at evaporative sites in leaves (VSMOW scale)
δin δ18O of CO2 in air entering leaf chamber (VSMOW scale)
δL δ18O of average lamina leaf water (VSMOW scale)
δR δ18O of CO2 efflux during dark respiration (VSMOW scale)
δs δ18O of source water (VSMOW scale)
E Transpiration rate
e Discrimination against 13C during day respiration
ea Vapor pressure in atmosphere
ei Vapor pressure in leaf intercellular air spaces
ɛ+ Equilibrium 18O fractionation between liquid water and vapor
ɛk Kinetic fractionation during diffusion of H218O from leaf intercellular air spaces to atmosphere
ɛw Equilibrium 18O fractionation between CO2 and water
f Discrimination against 13C during photorespiration
gt Conductance to H2O from leaf intercellular air space to atmosphere
gi Conductance to CO2 from leaf intercellular air spaces to sites of carboxylation
gtc Conductance to CO2 from chloroplast to atmosphere
Γ* CO2 compensation point in absence of day respiration
k Carboxylation efficiency
L Scaled effective path length for calculation of ℘
Λ Area of leaf in leaf chamber
m13 Slope of the relationship between Δi − Δobs and A/Ca
m Slope of the relationship between δc and δe
n Number of measurements in each experiment
Péclet number
θ Propotion of chloroplast CO2 isotopically equilibrated with chloroplast water
P Atmospheric pressure
RA 13C/12C or 18O/16O of net CO2 uptake by photosynthesis
Ra 13C/12C or 18O/16O of CO2 in atmosphere
Rce 18O/16O of CO2 in equilibrium with chloroplast water
Rc0 18O/16O of chloroplast CO2 that has not equilibrated with chloroplast water
Re 18O/16O of water at evaporative sites in leaves
RR 18O/16O of net CO2 efflux during dark respiration
RStd 18O/16O of VSMOW standard
rs Stomatal resistance to water vapor diffusion
rb Leaf boundary layer resistance to water vapor diffusion
d Day respiration rate
n Net CO2 efflux rate during dark respiration
ρ Ratio of rates of carboxylation and CO2 hydration in chloroplast
Tleaf Leaf temperature
u Flow rate of air through leaf chamber
wi Mole fraction of water vapor in the leaf intercellular air spaces
W Leaf lamina water concentration