A |
Net photosynthesis rate |
ā |
Weighted mean discrimination against C18OO for diffusion from chloroplast to atmosphere |
a |
Discrimination against C18OO during diffusion through stomata |
ab
|
Discrimination against C18OO during diffusion through leaf boundary layer |
aw
|
Summed discriminations against C18OO during liquid phase diffusion and dissolution |
a13
|
Discrimination against 13CO2 during diffusion through stomata |
ab13
|
Discrimination against 13CO2 during diffusion through leaf boundary layer |
aw13
|
Summed discrimination against 13CO2 during dissolution and liquid phase diffusion |
αw
|
Equilibrium oxygen isotope effect between CO2 and water |
b |
Discrimination against 13CO2 by carboxylating enzymes |
b18
|
Discrimination against C18OO by Rubisco |
C |
Molar concentration of water |
Ca
|
Partial pressure of CO2 in atmosphere |
Cc
|
Partial pressure of CO2 in chloroplast |
Ccs
|
Partial pressure of CO2 at the chloroplast surface |
Ci
|
Partial pressure of CO2 in leaf intercellular air spaces |
Cin
|
Partial pressure of CO2 in dry air entering leaf chamber |
Cs
|
Partial pressure of CO2 at the leaf surface |
ci
|
Mole fraction of CO2 in intercellular air spaces |
ca
|
Mole fraction of CO2 in atmosphere |
D |
Diffusivity of H218O in water |
ΔA
|
Discrimination against 13C or 18O during net CO2 uptake by photosynthesis |
Δca
|
18O enrichment of CO2 in chloroplast compared to atmosphere |
Δe
|
18O enrichment at evaporative sites in leaves compared to source water |
Δea
|
18O enrichment of CO2 in chloroplast compared to atmosphere when chloroplast CO2 is in full equilibrium with chloroplast water |
Δi
|
Discrimination against 13CO2 that would occur if gi were infinite and photorespiration and day respiration did not discriminate |
ΔL
|
18O enrichment of average lamina leaf water compared to source water |
Δobs
|
Observed discrimination against 13CO2 during photosynthesis |
Δv
|
18O enrichment of vapor in atmosphere compared to source water |
δA
|
δ18O of CO2 taken up by net photosynthesis (VSMOW scale) |
δa
|
δ18O of CO2 in atmosphere (VSMOW scale) |
δc
|
δ18O of CO2 in chloroplast (VSMOW scale) |
δc0
|
δ18O of CO2 in chloroplast that has not equilibrated with chloroplast water (VSMOW scale) |
δe
|
δ18O of water at evaporative sites in leaves (VSMOW scale) |
δin
|
δ18O of CO2 in air entering leaf chamber (VSMOW scale) |
δL
|
δ18O of average lamina leaf water (VSMOW scale) |
δR
|
δ18O of CO2 efflux during dark respiration (VSMOW scale) |
δs
|
δ18O of source water (VSMOW scale) |
E |
Transpiration rate |
e |
Discrimination against 13C during day respiration |
ea
|
Vapor pressure in atmosphere |
ei
|
Vapor pressure in leaf intercellular air spaces |
ɛ+
|
Equilibrium 18O fractionation between liquid water and vapor |
ɛk
|
Kinetic fractionation during diffusion of H218O from leaf intercellular air spaces to atmosphere |
ɛw
|
Equilibrium 18O fractionation between CO2 and water |
f |
Discrimination against 13C during photorespiration |
gt
|
Conductance to H2O from leaf intercellular air space to atmosphere |
gi
|
Conductance to CO2 from leaf intercellular air spaces to sites of carboxylation |
gtc
|
Conductance to CO2 from chloroplast to atmosphere |
Γ*
|
CO2 compensation point in absence of day respiration |
k |
Carboxylation efficiency |
L |
Scaled effective path length for calculation of ℘ |
Λ |
Area of leaf in leaf chamber |
m13
|
Slope of the relationship between Δi − Δobs and A/Ca
|
m |
Slope of the relationship between δc and δe
|
n |
Number of measurements in each experiment |
℘ |
Péclet number |
θ |
Propotion of chloroplast CO2 isotopically equilibrated with chloroplast water |
P |
Atmospheric pressure |
RA
|
13C/12C or 18O/16O of net CO2 uptake by photosynthesis |
Ra
|
13C/12C or 18O/16O of CO2 in atmosphere |
Rce
|
18O/16O of CO2 in equilibrium with chloroplast water |
Rc0
|
18O/16O of chloroplast CO2 that has not equilibrated with chloroplast water |
Re
|
18O/16O of water at evaporative sites in leaves |
RR
|
18O/16O of net CO2 efflux during dark respiration |
RStd
|
18O/16O of VSMOW standard |
rs
|
Stomatal resistance to water vapor diffusion |
rb
|
Leaf boundary layer resistance to water vapor diffusion |
ℜd
|
Day respiration rate |
ℜn
|
Net CO2 efflux rate during dark respiration |
ρ |
Ratio of rates of carboxylation and CO2 hydration in chloroplast |
Tleaf
|
Leaf temperature |
u |
Flow rate of air through leaf chamber |
wi
|
Mole fraction of water vapor in the leaf intercellular air spaces |
W |
Leaf lamina water concentration |