Table 1.

protein classes and their origins and function as derived in this study.

	First appearance	WGD	Function	Location	Reference
C-ter SUN	Moss		LINC complex component; binds KASH; Nuclear shape and size; meiosis	INM	Graumann et al.17; Oda and Fukuda, 2011
Mid-SUN	Alga		LINC complex component; binds KASH; Nuclear shape and size; fertility	INM and ER	Graumann et al.18
CRWN	Moss		nucleoskeleton; nuclear size and shape; heterochromatin organization	nuclear periphery and nucleoplasm	Dittmer et al.09; Wang et al., 2013
SINE1-2	Moss		LINC complex component; KASH; interacts with actin cytoskeleton; nuclear positioning in guard cells (SINE1); innate immunity response (SINE2)	ONM	Zhou et al.41,42
NEAP	Basal angiosperm, Gymnosperm	Zeta	NE anchor, SUN binding, chromatin interactor; root growth; nuclear morphology	INM	Pawar, 2015
WIP	Basal angiosperm	Epsilon	SUN binding; anchors RanGAP to NE; nuclear morphology; pollen tube termination; nuclear movement	ONM; RanGAP anchorage	Xu et al.38; Zhao et al., 2008
KAKU 4	Monocot	Epsilon	CRWN binding; nuclear size and shape	Nucleoskeleton	Goto et al.14
SINE 3-4	Eudicot (Brassicas)	Gamma	LINC complex component; KASH; SUN binding	ONM	Zhou et al.42
TIK	Eudicot (Arabidopsis)	Gamma	SUN binding; nuclear size and shape; root growth	ONM	Graumann et al.18