A Firing rates averaged across populations 5E and 6E and across areas for different stages from the original model (black) to iteration 4 (light gray) as a function of , predicted by the mean-field theory. B Relative changes in the indegree between areas A, B in the first iteration. C Layer-specific relative changes in the connections within and between areas FEF and 46, for the first iteration. Populations are ordered from 2/3E (left) to 6I (right) on the horizontal axis and from 6I (bottom) to 2/3E (top) on the vertical axis as in panel D. D Relative changes in population-specific indegrees summed over target populations, , combined for iterations two, three and four.