Microbial community biomass (a), richness (b), diversity (c), and evenness (d) after communities assembled under different dispersal levels (x-axis) and litter chemistries (lignin:N, symbols and colored lines) for six years. Species richness is the number of species present, while alpha diversity (measured by Shannon's diversity index) incorporates both richness and evenness. Taxa abundances were integrated over 365 days in year six and summed over the entire grid. Asterisks indicate a significant effect (P<0.01) of lignin:N, and error bars show standard error around the mean (N=6).