Table 1.

Sizes of species trees with different number of species generated under the protracted speciation process under different species conversion rates, c, the rate at which isolated lineages develop into true species

	Lineages			Species
c	Max	Min	Mean	Max	Min	Mean
0.001	47	3	17.35	2	1	1.01
0.1	38	4	13.29	7	1	1.83
1	36	3	11.43	12	1	3.75
10	39	3	10.43	17	1	4.87
1,000	39	3	10.67	19	1	5.46

At the lowest conversion rate (0.001, or 500 times slower than the speciation initiation rate), on average only one actual species was generated in each species phylogeny, even though the number of lineages ranged between 3 and 47. At the highest conversion rate (1,000 or 2,000 times faster than the speciation initiation rate), on average approximately five species were expected on phylogenies that ranged in size from 3 to 39 tips. Results here are based on a birth rate of 0.5 and extinction rates of 0.0 and 0.2 (see Materials and Methods for details).