In TSC2-deficient cells, activation of mTORC1 and Syk signaling induces MCP-1 overproduction via a Stat3-dependent manner. Subsequently, MCP-1 recruits and activates peripheral blood mononuclear cells (PBMCs), leading to increased VEGF-D expression. In contrast, treatment with Syk inhibitors impairs proliferation of TSC2-deficient cells and decreases production of MCP-1. As a result, VEGF-D expression is reduced in PBMCs.