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Modification positions in tRNA and the responsible SpoU-TrmD (SPOUT) tRNA methyltransferases. The secondary structure of tRNA is represented in cloverleaf structure. The conserved nucleotides in tRNA are depicted as follows: adenosine, A; guanosine, G; cytidine, C; uridine, U; purine, R; pseudouridine, Ψ. The modified positions are numbered and the associated enzymes are indicated in red. The structures of modified nucleosides and abbreviations of their names are shown in Figure 1.
