Overexpression of PIF3-nA4 in cop1–4 caused reduced photomorphogenic phenotype. Images (Upper) and angles of cotyledon opening (Lower) of dark-grown Col-0, PIF3-WT/Col-0, PIF3-nA4/Col-0, cop1–4, PIF3-WT/cop1–4, and PIF3-nA4/cop1–4 seedlings. Mean ± SD, n = 20. (Scale bar, 1 mm.) **P < 0.01 (t test).