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. 2017 Mar 1;12(3):e0171903. doi: 10.1371/journal.pone.0171903

Table 1. Range of sleep times and other relevant data reported for elephants previously in the literature.

Study Species Number and Sex Setting Total rest/sleep time (h) Timing of rest/sleep Timing of major sleep Time of awakening Standing rest/sleep observed? Recumbent rest/sleep observed?
Day Night
Benedict and Lee, 1938 [1] Asian and African Not reported Captive—Circus Not reported Yes, rarely and short duration Yes 00.00–04.30 04.30–05.00 Yes Yes
Kurt, 1960 [2] Asian (10) and African (7) 5 males, 12 females Captive—Circus 5.92 h No Yes 23.00–07.00 07.00 Yes Yes
Khüme, 1963 [3] African 2 males, 1 female Captive—Zoo 4.17 h Not reported Yes 03.00–05.00 05.00 Yes Yes
McKay, 1973 [4] Asian Both, many animals Wild–Sri Lanka Not reported Yes Not reported Not reported Not reported Yes Yes
Wyatt and Eltringham, 1974 [5] African 6 males, 12 females Wild—Uganda 3.17–3.36 h Yes, short duration Yes 04.00–07.00 07.00 Yes Yes
Guy, 1976 [6] African 27 males, 21 females Wild—Zimbabwe Not reported Yes, around 2.4 h Not reported Not reported Not reported Yes Not reported
Adams and Berg, 1980 [7] African 7 females Captive—Zoo Not reported Yes Not reported Not reported Not reported Not reported Yes
Tobler, 1992 [8] Asian 12 females Captive–zoo and circus 3.10–6.90 h Not reported Yes 01.00–04.00 06.00–07.00 Yes Yes
Brockett et al., 1999 [9] African 3 females Captive—Zoo Not reported Not reported Yes 01.00–06.00 06.00 Not reported Yes
Wilson et al., 2006 [10] African 3 females Captive—Zoo Not reported Not reported Yes 22.00–02.00 Not reported Yes Yes
Horback et al., 2014 [11] African 7 males, 8 females Captive—Zoo Not reported Yes Yes 23.00–04.00 05.00–06.00 Yes Yes
Holdgate et al., 2016 [12] Asian (28) and African (44) 72 females Captive—Zoo African > 2.1 h. Asian > 3.2 h Yes Yes 01:00–05:00 Not reported Yes Yes