Benedict and Lee, 1938 [1] |
Asian and African |
Not reported |
Captive—Circus |
Not reported |
Yes, rarely and short duration |
Yes |
00.00–04.30 |
04.30–05.00 |
Yes |
Yes |
Kurt, 1960 [2] |
Asian (10) and African (7) |
5 males, 12 females |
Captive—Circus |
5.92 h |
No |
Yes |
23.00–07.00 |
07.00 |
Yes |
Yes |
Khüme, 1963 [3] |
African |
2 males, 1 female |
Captive—Zoo |
4.17 h |
Not reported |
Yes |
03.00–05.00 |
05.00 |
Yes |
Yes |
McKay, 1973 [4] |
Asian |
Both, many animals |
Wild–Sri Lanka |
Not reported |
Yes |
Not reported |
Not reported |
Not reported |
Yes |
Yes |
Wyatt and Eltringham, 1974 [5] |
African |
6 males, 12 females |
Wild—Uganda |
3.17–3.36 h |
Yes, short duration |
Yes |
04.00–07.00 |
07.00 |
Yes |
Yes |
Guy, 1976 [6] |
African |
27 males, 21 females |
Wild—Zimbabwe |
Not reported |
Yes, around 2.4 h |
Not reported |
Not reported |
Not reported |
Yes |
Not reported |
Adams and Berg, 1980 [7] |
African |
7 females |
Captive—Zoo |
Not reported |
Yes |
Not reported |
Not reported |
Not reported |
Not reported |
Yes |
Tobler, 1992 [8] |
Asian |
12 females |
Captive–zoo and circus |
3.10–6.90 h |
Not reported |
Yes |
01.00–04.00 |
06.00–07.00 |
Yes |
Yes |
Brockett et al., 1999 [9] |
African |
3 females |
Captive—Zoo |
Not reported |
Not reported |
Yes |
01.00–06.00 |
06.00 |
Not reported |
Yes |
Wilson et al., 2006 [10] |
African |
3 females |
Captive—Zoo |
Not reported |
Not reported |
Yes |
22.00–02.00 |
Not reported |
Yes |
Yes |
Horback et al., 2014 [11] |
African |
7 males, 8 females |
Captive—Zoo |
Not reported |
Yes |
Yes |
23.00–04.00 |
05.00–06.00 |
Yes |
Yes |
Holdgate et al., 2016 [12] |
Asian (28) and African (44) |
72 females |
Captive—Zoo |
African > 2.1 h. Asian > 3.2 h |
Yes |
Yes |
01:00–05:00 |
Not reported |
Yes |
Yes |