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Average structures of the RE_94 (top left), UN_94 (top right) and OV_89 (bottom) minicircles, shown in a wire representation, surrounded by transparent surface envelopes. Each minicircle is oriented with base pair 1 toward the top of the figure, with the numbering increasing in a clockwise sense. Key features are the strongly unwound segment in UN_94 C29-A33 (indicated by the arrow) and the out-of-plane writhing of the overwound OV_89.
