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. 2017 Apr 21;7:1052. doi: 10.1038/s41598-017-01136-1

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© The Author(s) 2017

Open Access This article is licensed under a Creative Commons Attribution 4.0 International License, which permits use, sharing, adaptation, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons license, and indicate if changes were made. The images or other third party material in this article are included in the article’s Creative Commons license, unless indicated otherwise in a credit line to the material. If material is not included in the article’s Creative Commons license and your intended use is not permitted by statutory regulation or exceeds the permitted use, you will need to obtain permission directly from the copyright holder. To view a copy of this license, visit http://creativecommons.org/licenses/by/4.0/.

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Histological sections of interdigital skin of front paw in adult male (A), interdigital skin of front paw in yearling male (B), interdigital skin of hind paw in adult male (C), interdigital skin of hind paw in yearling male (D), ventral metatarsal skin in adult male (E), ventral metatarsal skin in yearling male (F), front footpad in adult male (G), front footpad in yearling male (H), and control lip skin (I) and control left shoulder skin (J) in adult male brown bear (Ursus arctos). In interdigital and metatarsal regions (A,B,C,D,E,F) there are hair follicles (HF) visible with apparently more profuse and prominent apocrine sweat glands (AG) in adult male (A,C and E). In sections of control skin of the adult male (I and J), apocrine sweat glands appear less profuse than in sections of the focus areas of the paws. Sections of footpads (G and H) show the stratified squamous keratinized epithelium of the pad with very thick striatum corneum (the darkest shade of stain), and eccrine glands (EG) deeper in the dermis of yearling male (H). Stain: hematoxylin and eosin. Magnification: x100. SG – sebaceous gland.