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. 2016 May 4;5(1):368. doi: 10.4102/ajlm.v5i1.368

TABLE 3.

Inducers of viable but non-culturable state in Escherichia coli.

Inducing factor Reference
Starvation 9, 11, 12, 16, 32, 33
Suboptimal temperature 12, 37
Chlorination 11, 16
Osmotic stress 43, 44
High pressure CO2 (HPCD) 13
Oxidative stress 10, 35, 40
Visible radiation 9, 32
Sunlight 45
pH variation 33