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Change in behaviours with female age. The relative amount of grooming females received (a) and the amount of aggression females gave (c) did not vary with age. By contrast, older adult females gave less grooming (b) and received less aggression (d) compared with younger adult females. Relative rates are z-scores, where values above 0 are above the mean rate of a given behaviour in a group in a given year, below 0 are below the mean in a group in a given year.
