External deformations on the fish caused by the bacteria. Hemorrhages on around the mouth (A), and the base of dorsal and caudal fins (B, C), unilateral exophthalmus (black arrow) on the fish Acipenser gueldenstaedtii, spiraculum (white arrow) (D) (scale bars, A: 2 cm, B: 2 cm, C: 5 cm, and D: 5 cm