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Effects of environmental setting (a) on the maximum depth of sediment particle reworking (f-SPILmax, cm, mean ± s.e., n = 3) for H. diversicolor (triangles) and H. ulvae (circles), and the effect of population (b) for H. ulvae (circles). Observations without macrofauna (dash, n = 9) are shown for comparison. The dotted line indicates the sediment surface and negative values indicate deeper net downward transport of sediment particles.
