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. 2017 Apr 18;58(6):1044–1054. doi: 10.1194/jlr.R073452

TABLE 2.

Frequency of appearance of the proposed common structural features among cholesterol/cholesterol sulfate binding sites on crystallized soluble proteins

Feature of Steroid-Binding Protein Site PDB Entry Frequency of Appearance
1LRI 1ZHY 4BOE 3GKI 3N9Y 5L7D 2HKA 1S0X 2Q9F
Protein pocket Yes Yes Yes Yes Yes Yes Yes Yes Yes 9/9
Avoidance of secondary structure-lacking areas Yes Yes Yes Yes Yes Yes Yes Yes Yes 9/9
Steroid α-face facing an α-helix Yes Yes Yes Yes Yes Yes 6/9
Steroid β-face facing a β-strand Yes Yes Yes Yes Yes Yes 6/9
Hydrogen-bonding with Asn, Gln, or Tyr Yes Yes Yes Yes Yes 5/9
Hydrogen bonding amino acid is on an α-helix Yes Yes Yes Yes Yes 5/9
Hydrophobic interactions with Leu, Ile, Val, or Phe Yes Yes Yes Yes Yes Yes Yes Yes Yes 9/9
Steroid-protein contacts at C21 and C26 Yes Yes Yes Yes Yes 5/9
Hydrogen bonding with O at C3 moiety Yes Yes Yes Yes Yes Yes Yes 7/9
Lack of contacts with the protein at steroid C8–C10, C13, and C17 Yes Yes Yes Yes Yes Yes Yes Yes Yes 9/9