VirE3 NLS1 and NLS2 promote nuclear import of GUS reporter in plant cells: (A, B) GUS-NLS1; (C, D) GUS-mNLS1; (E, F) GUS-NLS2; and (G, H) GUS-mNLS2. Panels A, C, E, and G show GUS staining, and panels B, D, F, and H show DAPI staining. Arrows indicate cell nuclei.