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. 2017 May 30;8:15696. doi: 10.1038/ncomms15696

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Copyright © 2017, The Author(s)

Open Access This article is licensed under a Creative Commons Attribution 4.0 International License, which permits use, sharing, adaptation, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons license, and indicate if changes were made. The images or other third party material in this article are included in the article's Creative Commons license, unless indicated otherwise in a credit line to the material. If material is not included in the article's Creative Commons license and your intended use is not permitted by statutory regulation or exceeds the permitted use, you will need to obtain permission directly from the copyright holder. To view a copy of this license, visit http://creativecommons.org/licenses/by/4.0/

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Under the drier scenario (a) hominin survival is focused on single springs (or spring clusters) and movement between springs (or spring clusters) is limited. As climate improves (b) the availability of water sources increases particularly as the water table intersects rift flank rivers. Springs high on the rift sides may act to link rift flank rivers with water sources in the rift facilitating transverse rift movement. As water becomes widely available (c) hominin movement occurs in all directions including along the rift axis.