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. 2017 Jun 7;7:3007. doi: 10.1038/s41598-017-03020-4

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© The Author(s) 2017

Open Access This article is licensed under a Creative Commons Attribution 4.0 International License, which permits use, sharing, adaptation, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons license, and indicate if changes were made. The images or other third party material in this article are included in the article’s Creative Commons license, unless indicated otherwise in a credit line to the material. If material is not included in the article’s Creative Commons license and your intended use is not permitted by statutory regulation or exceeds the permitted use, you will need to obtain permission directly from the copyright holder. To view a copy of this license, visit http://creativecommons.org/licenses/by/4.0/.

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M. marinum ΔwhiB4 exhibited altered colony morphology and was defective in cording formation. (A) M. marinum ΔwhiB4 exhibited dry and rough colony morphology on 7H11 agar, as compared to as compared to the smoother, glossier, morphology of WT or the complemented strain. (B) Cells of WT or the complemented strain formed the serpentine cords, which were not observed for ΔwhiB4.