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. 2017 Jun 13;7:3428. doi: 10.1038/s41598-017-03393-6

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© The Author(s) 2017

Open Access This article is licensed under a Creative Commons Attribution 4.0 International License, which permits use, sharing, adaptation, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons license, and indicate if changes were made. The images or other third party material in this article are included in the article’s Creative Commons license, unless indicated otherwise in a credit line to the material. If material is not included in the article’s Creative Commons license and your intended use is not permitted by statutory regulation or exceeds the permitted use, you will need to obtain permission directly from the copyright holder. To view a copy of this license, visit http://creativecommons.org/licenses/by/4.0/.

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Changes in the amount of LMW-GSs in IG and SG in the six deletion mutants cultivated in five environments. Xiaoyan 81 and the six deletion lines (DLGluA1, DLGluB1, DLGluD1, DLGluA1B1, DLGluA1D1 and DLGluB1D1) were grown in two locations (ZX and XX) in 2013/2014 and three locations (BJ, ZX and XX) in 2014/2015. The resultant grain samples were analyzed for the amounts of LMW-GSs in the IG and SG fractions, respectively. The values shown are means ± SE of three separate tests, with that of Xiaoyan 81 being set as 1 to facilitate comparisons. In each environment, the means marked by different letters are statistically significant (P < 0.05). The amounts of LMW-GSs in IG were generally decreased in the six deletion lines compared to that in Xiaoyan 81 (A). On the contrary, the amounts of LMW-GSs in SG were generally increased in the six deletion lines relative to that in Xiaoyan 81 (B). BJ, Beijing; IG, insoluble glutenin; LMW-GSs, low-molecular-weight glutenin subunits; SG, soluble glutenin; XX, Xinxiang; ZX, Zhaoxian.