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Mass spectrometry identified human conventional and ancestral human H4 histone variants. Direct analyses and narrow scan in mass range 730–740 m/z; MSMS for center mass 734.40 m/z (2 nd isotope of 733.9 m/z; z=2); MSMS for center mass 744.40 m/z (2 nd isotope of 734.9 m/z; z=2) confirmed the co-existence of the human and ancestral H4 isoforms (with the IVY pattern) at 734.905 m/z; z=2.
