Table 2. The environmental predictor sets used in the different models.

Alternative predictor sets used in models.

Abbreviation	Environment variable	VC	VC+S	VS
T	Annual mean temperature	•	•
TM warm	Maximum temperature of warmest month	•	•
P	Mean annual precipitation	•	•
PQ warm	Precipitation of warmest quarter	•	•
PQ cold	Precipitation of coldest quarter	•	•
BD	Bulk soil density (g/cm3)		•	•
CLAY	Clay content percentage		•	•
pH	pH CaCl2		•	•
OC	Organic carbon percentage		•	•

Notes.

V_C

climate variables only

V_C+S

climate and soil variables

V_S

soil variables only