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. 2017 Jun 27;7:4281. doi: 10.1038/s41598-017-04368-3

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© The Author(s) 2017

Open Access This article is licensed under a Creative Commons Attribution 4.0 International License, which permits use, sharing, adaptation, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons license, and indicate if changes were made. The images or other third party material in this article are included in the article’s Creative Commons license, unless indicated otherwise in a credit line to the material. If material is not included in the article’s Creative Commons license and your intended use is not permitted by statutory regulation or exceeds the permitted use, you will need to obtain permission directly from the copyright holder. To view a copy of this license, visit http://creativecommons.org/licenses/by/4.0/.

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Expression and correlation of USP3, SMAD4 and miR-224 in primary CRC samples (a–c) USP3, SMAD4 and miR-224 expression in 40 paired primary CRC and adjacent non-cancer tissues were evaluated by qRT-PCR. (d) Comparison of the USP3 expression levels between “SMAD4 low” and “SMAD4 high” subgroups that were classified according to the median SMAD4 expression levels. (e–g) Correlation analyses of USP3 mRNA, SMAD4 mRNA and miR-224 were conducted in these CRC specimens.