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. 2017 Jun 30;7:4477. doi: 10.1038/s41598-017-04750-1

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© The Author(s) 2017

Open Access This article is licensed under a Creative Commons Attribution 4.0 International License, which permits use, sharing, adaptation, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons license, and indicate if changes were made. The images or other third party material in this article are included in the article’s Creative Commons license, unless indicated otherwise in a credit line to the material. If material is not included in the article’s Creative Commons license and your intended use is not permitted by statutory regulation or exceeds the permitted use, you will need to obtain permission directly from the copyright holder. To view a copy of this license, visit http://creativecommons.org/licenses/by/4.0/.

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Unigene expression involved in shikonin biosynthesis in Lithospermum erythrorhizon, Arnebia euchroma and Echium plantagineum. (a) Shikonin biosynthesis; (b) Unigene expressions involved in shikonin biosynthesis? One step unknown?* Several steps unknown; 1. Acetoacetyl-coenzyme A thiolase gene (AACT), 2. Hydroxy-methylglutaryl CoA synthase gene (HMGS), 3. Hydroxy-methylglutaryl-CoA reductase gene (HMGR), 4. Mevalonate kinase gene (MK), 5. Phosphomevalonate kinase gene (PMK), 6. Mevalonate disphosphate decarboxylase gene (MVD), 7. Geranyl diphosphate synthase gene (GDS), 8. Phenylalanine ammonia-lyase gene (PAL), 9. 4-Coumarate:CoA ligase gene (4CL), 10. Cinnamic acid 4-hydroxylase gene (C4H), 11. p-Hydroxybenzoate geranyltransferase gene (PGT), 12. Geraniol 10-hydroxylase gene (G10H, predicted).