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. 2017 Jul 3;372(1727):20160238. doi: 10.1098/rstb.2016.0238

Table 1.

Model parameters and variables, their notation, level of operation and values.

variable/parameter notation level description value
nutritional state NS individual an individual's nutritional state, as tracked by its (p, c) position in the nutrient space, and denoting total intake of the two nutrients variable (p, c)
intake target IT global an individual's IT is the (p, c) coordinate in the nutrient space that maximizes F variable (p, c)
distance to the IT D individual the Euclidean distance between an individual's NS and the IT variable
food rail V global the nutritional composition of a food in terms of the amount of carbohydrate (c) in a given food relative to single equivalent unit of protein (p) 1, 3, 0.333, 16 and 0.667 (figure 1)
appetite A individual the amount of a given food that an individual would consume to minimize D variable
ideal food rail αideal individual the angle of a hypothetical ideal food rail connecting an individual's NS with their IT variable
meal size φ global the maximum amount of food that an individual is able to consume at a given time 0.0368
social influence σ individual the weight of the social interactions relative to the individual orientation when moving 0, 0.005, 0.01, 0.025, 0.2
noise weight η individual the weight of a random unit vector relative to vectors corresponding to social and individual influences during movement fixed at 0.0025, 0.0275 or 0.0525; or variable (see §2a(iii) Movement rules)
higher noise limit U individual upper bound value when calculating the variable noise as a function of D 0.0525
lower noise limit L individual lower bound value when calculating the variable noise as a function of D 0.0025
noise decrease constant K individual slope constant when calculating the variable noise as a function of D 0.0075
repulsion range Rr individual range at which neighbours exert a repulsion force on an individual 1
alignment range Ra individual range at which an individual aligns with its neighbours 6
attraction range Rp individual range at which an individual is attracted to its neighbours 56
proportion of individuals within alignment range psoc psoc global the proportion of moving individuals who have at least one neighbour with alignment range Ra or closer variable
spatial coordinates of an individual Λ individual the {x, y} spatial coordinates of an individual variable