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. 2017 Jul 11;7:5137. doi: 10.1038/s41598-017-04973-2

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© The Author(s) 2017

Open Access This article is licensed under a Creative Commons Attribution 4.0 International License, which permits use, sharing, adaptation, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons license, and indicate if changes were made. The images or other third party material in this article are included in the article’s Creative Commons license, unless indicated otherwise in a credit line to the material. If material is not included in the article’s Creative Commons license and your intended use is not permitted by statutory regulation or exceeds the permitted use, you will need to obtain permission directly from the copyright holder. To view a copy of this license, visit http://creativecommons.org/licenses/by/4.0/.

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Simultaneous chemical and sensory analyses of compounds and scents in lion MF headspace. Top seven most odorous compounds in lion marking fluid based on measured odor intensity. Chromatogram (top) highlighting identified compounds in lion MF do not necessarily follow their measured odor intensity (aromagram, bottom). The odor character descriptions are based on panelists’ evaluations. Aromagram was created by panelists during sensory analyses, recording odor character, intensity and start-end detection times.