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. 2017 Jul 13;7:5324. doi: 10.1038/s41598-017-05641-1

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© The Author(s) 2017

Open Access This article is licensed under a Creative Commons Attribution 4.0 International License, which permits use, sharing, adaptation, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons license, and indicate if changes were made. The images or other third party material in this article are included in the article’s Creative Commons license, unless indicated otherwise in a credit line to the material. If material is not included in the article’s Creative Commons license and your intended use is not permitted by statutory regulation or exceeds the permitted use, you will need to obtain permission directly from the copyright holder. To view a copy of this license, visit http://creativecommons.org/licenses/by/4.0/.

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Representative MALDI-TOF-MS spectra of EPO N-glycans. (A) A spectrum of the EPO standard. It should be noted that the antenna isomerism indicated for the partially sialylated tri-antennary glycan also applies to all other tri-antennary structures. In addition, there may be minor contributions of bi-antennary structures with one LacNAc repeat. The tetra-antennary glycans have significant contributions of tri-antennary structures with one additional LacNAc repeat, as exemplified for the glycan with two NANA and 4 LacNAc units. (B) The major N-glycans detected for EPO purified from CCS were very similar to those of the EPO standard albeit in different relative abundances. 2× and 3× gives the incidence of the monosaccharide or antenna to the right or below. The brackets signify that the exact positions of substituents are unclear. The N-acetylneuraminic acids to the right of the vertical brackets may cap an antenna or LacNAc-repeat.