Table 2. Evolutionary timescales of FVs and FLERVs.

Node*	Total per-lineage substitution (substitutions per site)		Evolutionary timescale (million years)†		Ref
	Median	95% HPD‡	Median	95% HPD
I	0.031	0.017–0.048	0.96	0.69–1.21	(ref. 39)
II	0.080	0.057–0.107	8.29	6.52–10.05	(ref. 40)
III	0.126	0.097–0.163	16.52	13.33–19.61
IV	0.160	0.123–0.199	31.60	25.36–37.85
V	0.292	0.238–0.349	43.43	38.39–48.08
VI	0.348	0.288–0.413	86.92	76.34–97.22
VII	0.381	0.315–0.454	98.59	95.94–100.78	(ref. 35)
VIII	0.436	0.357–0.523	100.95	98.40–103.83
IX	0.559	0.464–0.660	103.92	99.73–109.09
X	0.764	0.641–0.889	262.76	195.00–342.08	NA
XI	0.908	0.771–1.056	347.89	250.62–479.97
XII	1.071	0.869–1.274	454.65	303.69–683.66

^*Referring to the node numbers in the viral tree in Fig. 3 (left). Total per-lineage substitution and evolutionary timescale estimates of nodes I–IX were used to estimate the power-law time-dependent rate phenomenon model (Fig. 4). The model was then extrapolated to infer the timescale of nodes X–XII from their total per-lineage substitution estimates.

^†Evolutionary timescale of nodes I–IX were directly inferred from those of their hosts. The references refer to the host date references.

^‡‘95% HPD’=95% highest probability density interval.