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Tylosin biosynthetic gene cluster of S. fradiae. Five regulatory genes are shown: TylP is a GBL signal receptor. TylQ is a transcriptional repressor and blocks tylosin biosynthesis by controlling the expression of tylR. tylT and tylS encode cluster-situated regulatory proteins of the SARPs family. tylU is a positive regulatory gene of tylosin biosynthesis. TylR is also a CSR
