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. 2017 Jul 31;8:867. doi: 10.3389/fimmu.2017.00867

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Copyright © 2017 Beirnaert, Desmyter, Spinelli, Lauwereys, Aarden, Dreier, Loris, Silence, Pollet, Cambillau and de Haard.

This is an open-access article distributed under the terms of the Creative Commons Attribution License (CC BY). The use, distribution or reproduction in other forums is permitted, provided the original author(s) or licensor are credited and that the original publication in this journal is cited, in accordance with accepted academic practice. No use, distribution or reproduction is permitted which does not comply with these terms.

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Three-dimensional structures of the three tumor necrosis factor (TNF)–Nanobody™ complexes. (A) Complex TNF–VHH#1; TNF and VHH#1 surfaces are colored orange and red, respectively. Shown right: details of the interaction of VHH#1 with TNF; VHH#1 in ribbon representation. (B) Complex TNF–VHH#2; TNF and VHH#2 surfaces are colored orange and green, respectively. (C) Complex TNF–VHH#3; TNF and VHH#3 surfaces are colored orange and blue, respectively.