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. 2017 Jul 31;7:6951. doi: 10.1038/s41598-017-07534-9

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© The Author(s) 2017

Open Access This article is licensed under a Creative Commons Attribution 4.0 International License, which permits use, sharing, adaptation, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons license, and indicate if changes were made. The images or other third party material in this article are included in the article’s Creative Commons license, unless indicated otherwise in a credit line to the material. If material is not included in the article’s Creative Commons license and your intended use is not permitted by statutory regulation or exceeds the permitted use, you will need to obtain permission directly from the copyright holder. To view a copy of this license, visit http://creativecommons.org/licenses/by/4.0/.

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AOH disrupted the cytoskeletal (F-actin) organization. F-actin microfilaments were contiguous along the apical cell border and formed a circumferential, double-banded structure under control conditions. Rapidly after AOH, the double-banded structure was lost and F-actin exhibited a diffuse distribution. Two days later, the distribution of F-actin was not limited to the cell border but instead was diffusely scattered throughout the cytoplasm. The F-actin distribution only partially recovered and its double-banded structure reappeared in some cells at Day 7.