Task-related changes in rnoise enhance coding in A1, not in ML. Values above 0 indicate that rnoise in the active condition uniquely brings about an improvement in pairwise performance. rnoise changes in ML (A) do not uniquely contribute to the observed increase in pairwise performance shown in Fig. 3. However, in A1 (B), rnoise does make a contribution to increased coding accuracy. In ML (C and E), task-related rnoise effects do not contribute to average performance differences in either the positive (C) or negative (E) rtuning groups. However, at AM depths of 60 and 80%, rnoise contributes to decreased (C) or increased (E) performance, in pairs with positive or negative rtuning, respectively. In A1 (D and F), task-related rnoise effects appear to enhance sensory coding across both rtuning categories.