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. 2017 Aug 3;7:7248. doi: 10.1038/s41598-017-07476-2

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© The Author(s) 2017

Open Access This article is licensed under a Creative Commons Attribution 4.0 International License, which permits use, sharing, adaptation, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons license, and indicate if changes were made. The images or other third party material in this article are included in the article’s Creative Commons license, unless indicated otherwise in a credit line to the material. If material is not included in the article’s Creative Commons license and your intended use is not permitted by statutory regulation or exceeds the permitted use, you will need to obtain permission directly from the copyright holder. To view a copy of this license, visit http://creativecommons.org/licenses/by/4.0/.

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(a) The original IDH proposition made by Connell¹. The author¹ considered diversity of a given community to be the highest at intermediate disturbance because of co-occurrence of colonist and climax species in the middle of disturbance gradient. From Connell¹ we derived theoretically that expected diversity of (b) Colonist species fraction peaks at high disturbance and that of (c) Climax species fraction peaks at low disturbance.