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. 2017 Aug 9;12(8):e0181504. doi: 10.1371/journal.pone.0181504
1. Clitellum annular evenly developed also on the ventral side 2
Clitellum saddle shaped, ventrally always less developed 3
2. Male pore postsetal, clitellum on 23-28 B. palustris Moore, 1895
Male pore setal on a huge porophore, clitellum ends on 30 B. schwerti sp. nov.
3. Clitellum ends on or after segment 34 4
Clitellum ends on or before segment 33 6
4. Clitellum on 24, 25-34, 35 5
Clitellum on 27-37 B. zeteki (Smith & Gittins, 1915)
5. Septa 8/9-11/12 thickened, setal ratio ab:cd = 1.5:1 B. welchi (Smith, 1917)
Septa 12/13-14/15 thickened, setal ratio ab:cd = 1:1 B. lawrenceae Fender, 1994
6. Clitellum ends before segment 31 7
Clitellum ends on or after 31 9
7. Septa 7/8-9/10 thickened, typhlosole bifid B. gieseleri (Ude, 1895)
All septa membranous, Typhlosole lamellar 8
8. Clitellum on 24 (25) -30, color dark brown-red B. parvus (Eisen, 1874)
Clitellum on ½21, 22-29, ½30, color light red-violet B. tumidus (Eisen, 1874)
9. Septa strongly 7/8-11/12 thickened, clitellum on 22, 23-32, 33 B. longicinctus (Smith & Gittins, 1915)
All septa membranous or just slightly thickened 10
10. Prostomium tanylobic B. eiseni (Levinsen, 1884)
Prostomium epilobic 11
11. Clitellum on ½24, 25-32, ½33 B. heimburgeri (Smith, 1928)
Clitellum on 26-31, ½23 B. rubidus (Savigny, 1826)