Fig. 8.

Serial processing of whisking signals within the cerebellar cortex. Voluntary movement (orange trace; lower box) is encoded by GC populations via linear increases in spike output (red trace; upper box) to represent whisker position. Excitation from GCs (via PFs) drives elevated firing rates in the molecular layer during whisker movement. Local inhibitory networks mediate sign-reversed responses (decreasing activity during whisking) in neighbouring INs (green trace; upper box). GCL granule cell layer, ML molecular layer, MI molecular layer interneuron, PF parallel fibre