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. 2017 Aug 17;7:8527. doi: 10.1038/s41598-017-08938-3

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© The Author(s) 2017

Open Access This article is licensed under a Creative Commons Attribution 4.0 International License, which permits use, sharing, adaptation, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons license, and indicate if changes were made. The images or other third party material in this article are included in the article’s Creative Commons license, unless indicated otherwise in a credit line to the material. If material is not included in the article’s Creative Commons license and your intended use is not permitted by statutory regulation or exceeds the permitted use, you will need to obtain permission directly from the copyright holder. To view a copy of this license, visit http://creativecommons.org/licenses/by/4.0/.

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After collisions between faecal pellets in the distal colon, the mucus layers covering their surface can be internalised into the resulting composite pellet. (a) AB/H/E stained longitudinal section of distal mouse colon. (b) Close-up showing the internalised firm mucus layer. (c) AB/H/E stained longitudinal section of distal mouse colon. A firm mucus layer devoid of bacteria is found inside the pellet. (d) FISH stained (Bacteria in red; nuclei in blue) longitudinal section of mouse distal colon, the internalised firm mucus layer is recognised by the absence of bacteria.