Table 2.

A table of definitions and constraints for each parameter and variable used within the model.

Symbol	Description	Constraint/Expression
$M_i^t$	Proportion of males of genotype i in	${\sum }_{i=1}^9(M_i^t+F_i^t)=1$
	generation t (normalised)	and $0\le M_i^t\le 1$
$F_i^t$	Proportion of females of genotype i in	${\sum }_{i=1}^9(M_i^t+F_i^t)=1$
	generation t (normalised)	and $0\le F_i^t\le 1$
$M_i^e$	Proportion of males of genotype i in	–
	generation t
$F_i^e$	Proportion of females of genotype i in	–
	generation t
$\overline{\Omega }$	Average fitness of whole population in the	${\sum }_{i=1}^9(M_i^e+F_i^e)$
	current generation
ϵA	Relative fitness of individuals with one copy	0 ≤ ϵA ≤ 1
	of construct A
ϵB	Relative fitness of individuals with one copy	0 ≤ ϵB ≤ 1
	of construct B
${\eta }_i^A$	Number of copies of construct A carried by an	${\eta }_i^A=0,1,2$
	individual of genotype i
${\eta }_i^B$	Number of copies of construct B carried by an	${\eta }_i^B=0,1,2$
	individual of genotype i
${\gamma }_i^{M,F}$	Lethality to males (M) or females (F)	$0\le {\gamma }_i^{M,F}\le 1$
	due to presence of constructs in individuals
	of genotype i
${\Omega }_i^{M,F}$	Total relative fitness of males (M) or	${\Omega }_i^{M,F}={ϵ}_A^{{\eta }_i^A}{ϵ}_B^{{\eta }_i^B}(1-{\gamma }_i^{M,F})$
	females (F) of genotype i possessing
	copies of the constructs, including lethality
	in the current generation
α	Release ratio (α:1, introduced:wild)	$\alpha =$Introduced/Wild-Type