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. 2017 Aug 21;7:8987. doi: 10.1038/s41598-017-09344-5

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© The Author(s) 2017

Open Access This article is licensed under a Creative Commons Attribution 4.0 International License, which permits use, sharing, adaptation, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons license, and indicate if changes were made. The images or other third party material in this article are included in the article’s Creative Commons license, unless indicated otherwise in a credit line to the material. If material is not included in the article’s Creative Commons license and your intended use is not permitted by statutory regulation or exceeds the permitted use, you will need to obtain permission directly from the copyright holder. To view a copy of this license, visit http://creativecommons.org/licenses/by/4.0/.

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Skeletal architecture of L. pertusa. (a) Sketch showing a typical branch of L. pertusa formed by a succession of corallites. Light (b) and electron (c) micrographs of the area of investigation. (d) Reflected-light image of the area highlighted in (c) displaying a thick rapid accretion deposit (RAD) area running horizontally across the view, and a thinner RAD lamella in the internal part of the corallite wall. Smaller roughly equidimensional RADs (not shown, s. Fig. 2) are also present within the thickening deposit (TD). Alternation of opaque (O) and translucent (T) bands can be distinguished in the TD area. Radial and longitudinal growth directions are indicated by R and L, respectively. Artwork 1a by A. Lethiers, used with permission.