Table 1.

Symbols

Cell Surface Tensions
β	cortical tension at aggregate/tissue surfacea
β∗	residual tension at cell-cell contacts, per cella
βc	cortical tension at cell-cell contacts per cell
βi	cell surface tension at gap
βg	cortical tension at gap
Γ	adhesion tension at cell-cell contacts
Γg	adhesion tension at gaps
Adhesion Strengths
σ	tissue surface tension: absolute adhesion strengtha
σi	internal surface tension, adhesion strength at gaps
α	relative adhesiveness
αcrit	critical relative adhesiveness for gap closure
αi	internal relative adhesiveness at gaps
Pressures
pc	cell hydrostatic pressure
pi	interstitial pressure
Geometry
r	radius of circle circumscribing the gap
rc	radius of cell-cell contact area
R	radius of curvature
Rs	radius of cell surface curvature at tissue surface
Ri	radius of cell surface curvature at gap
l	gap size
θ	contact angle at tissue surface
θi	contact angle at gap

^aRefers to values from previous work (1).