Table S3.

Sample size of observed dataset

Observed dataset	n
Total included in sample	98
Placed as guide or breeder	66
Released from program for behavioral reasons	32
Total excluded from sample	40
Transferred to external organization	1
Died	1
Released from program for medical reasons	26
Missing novel object data due to camera malfunction	1
Missing multistep problem-solving data due to failing to pass the warm-up trials	8
Missing all young adult data due to release before return to headquarters	3