Endemicity boundaries in the presence of periodically forced transmission (model (3)). Long-term pathogen establishment is possible () on the right of the endemicity curves, which correspond to . A) Periodic veriations of exposure/transmission risk (black: and ) or of the population abundance of either host species (gray: and either or ). B) Coupled fluctuations of exposure/transmission risk and of the population abundance of either host species (black: and either and or and ), or of the population abundances of the two host species (gray: and ), for different phase shifts between the relevant periodic signals (labels). C) As in panel B, for synchronous coupled fluctuations with different amplitudes (black: and either and or and ; gray: and ). Results are shown for different values of the basic homogeneous reproduction number r0 (labels). In all panels, ; other parameters as in Fig. 1.