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. 2017 Sep 26;7:12329. doi: 10.1038/s41598-017-12190-0

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© The Author(s) 2017

Open Access This article is licensed under a Creative Commons Attribution 4.0 International License, which permits use, sharing, adaptation, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons license, and indicate if changes were made. The images or other third party material in this article are included in the article’s Creative Commons license, unless indicated otherwise in a credit line to the material. If material is not included in the article’s Creative Commons license and your intended use is not permitted by statutory regulation or exceeds the permitted use, you will need to obtain permission directly from the copyright holder. To view a copy of this license, visit http://creativecommons.org/licenses/by/4.0/.

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Indentation of C. elegans. (A) Measurement of displacement and applied force as a function of time for an adult worm indented around its mid-body (B) Typical brightfield images captured at 0 µm (before) and at 12 µm (after) indentation. (C) Deconvolved fluorescence images showing deformation of the DiI labelled cuticle at different indentation depths (displayed as inverted grayscale). (D) 3D isosurface (rendered from deconvolved fluorescence images) showing C. elegans cuticle before (left) and after (right) indentation to a depth of 12 µm (right) illustrating the extent and topography of the resulting deformation.