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MERS-CoV S1-mediated Sia-binding activity resides in S1A. MERS-CoV S1-, S1A-, and S1B-Fc proteins alone or complexed with pA-LS were subjected to hemagglutination assay using human erythrocytes, after which HA titers were determined. MERS-CoV S1A-Fc hemagglutination activity was additionally tested with NA-treated human erythrocytes. Wells positive for hemagglutination are encircled. Hemagglutination assays were performed at least two times.
