Fig 9. Coefficient of variation (CoV; standard deviation/mean*100%) of the half-sarcomere lengths during an isometric contraction at muscle length L/L₀ = 1.2, for models ${\text{M}}_{\text{ATI}}^{\text{Fv}}$ (blue line), M^Fv (red line), M_ATI (black solid line) and ${\text{M}}_{\text{x}}^{\text{x}}$ (black dashed line).

While actin—titin interactions can damp the magnitude of half-sarcomere—length heterogeneities, the force-velocity relation damps the their temporal evolution.