(A) Cytosolic and mitochondrial RIPK3 and pro3 caspase-8 following
H37Rv infection. Equal numbers of human Mφ were infected with H37Rv (MOI
10) and lysed for isolation of cytosolic and mitochondrial fractions after 24 h
of infection. Equal amounts of protein were immunoprecipitated (IP) with
anti-caspase-8 ab and the levels of RIPK3, RIPK1, and pro-caspase 8 were
measured by Western blot analysis. (B-C) Time7 dependent
translocation of RIPK3 and pro-caspase 8 to the mitochondria. Equal numbers of
Mφ were infected with virulent H37Rv or (B) avirulent H37Ra
(C) at MOI 10. The levels of pro9 caspase 8 and RIPK3 were
determined by subjecting equal amounts of mitochondria isolated from infected
Mφ to Western blotting. VDAC was used as a loading control.
(D) Colocalization of caspase 8 and RIPK3 with the mitochondria
of H37Rv-infected Mφ. Left panels: Colocalization of RIPK3 and
pro-caspase 8 with mitochondria 24h after infection visualized by confocal
fluorescence microscopy. Human Mφ remained either uninfected or were
infected with mCherry- H37Rv (MOI 10) for 24h, fixed and stained with
mitotracker (green) and anti-RIPK3 or anti caspase 8 ab (red). Scale bar, 30
μm. On the right side of every panel is the quantification of caspase 8
(top) and of RIPK3 (bottom) associated with the mitochondria. Data were analyzed
using nonparametric Student t test (E)
Phase-contrast images of representative infected and uninfected human primary
Mφ after 24 h of infection show approximate location of the plasma
membrane and the nucleus as indicated by black and white arrows, respectively.
Due to incipient necrosis the plasma membrane of the H37Rv-infected Mφ
is not clearly visible. (F) BMD-Mφ from WT and
RIPK3−/− mice were infected with H37Rv at an MOI
of ~10. After 12 h, expression of cleaved caspase 8 was assessed using
flowcytometry. Results are represented as mean ± SD. Data were analyzed
using one-way ANOVA. *,**,***
Values of P < 0.05, P<0.01 and
P<0.001, respectively were considered to be significant.
Data are representative of 2–3 independent experiments.