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Initiation of RT when (A) wt-β2 or (B) XY-β2 (where X is NO2 or Fn) reacts with α2, substrate CDP (S) and allosteric effector ATP (E). (A) Reduction of wt Y122–O• by Y356 is coupled with proton transfer (PT) from Fe1-H2O.64 (B) Reduction of NO2Y122–O• (or FnY122–O•) by Y356 is not coupled to PT from the diiron cluster.
