Genome organization and translation strategy of beet necrotic yellow vein virus. The scheme indicates self-cleavage of the replicase protein (red arrow and black triangle), a suppressible UAG stop codon (green star), m7Gppp (black circle) and the 3′ poly(A) tails (An). Mtr, methyltransferase; Hel, helicase; Pro, protease; Pol, RNA polymerase; RT, readthrough; RTD, readthrough domain; sub, subgenomic; CRP, cysteine rich protein. N and p4.6 have never been detected. Noncoding RNAs, ncRNA3 and ncRNA5, produced by exoribonuclease activity harbour the conserved coremin motif present in the ‘core region’ responsible for long distance movement in Beta species.